Main Ref. | SAUP Database, 2006 |
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Remarks | Coastal to <10 km offshore (Ref. 356). Found on coastal rock, and over sea (Ref. 83946). Breeds annually in dense colonies, preferably in bushes and trees in Atlantic and Pacific Oceans (Ref. 84934). They are double-brooded. When there is danger, alarm calls are given by adults and chicks respond by crouching in place (Ref. 87784). Highly pelagic. Typically feeds far out at sea in large, gregarious flocks over shoals of predatory fish (Ref. 91362). Exhibits dipping and surface-seizing foraging behavior (Ref. 356). Feed mainly by dipping, hovering and picking from or just below the surface of the water. Also, shallow-plunging (Ref. 126735). |
Marine - Neritic | Marine - Oceanic | Brackishwater | Freshwater | |
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Marine zones / Brackish and freshwater bodies |
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Substrate | Hard Bottom: rocky; |
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Substrate Ref. | |
Special habitats | |
Special habitats Ref. |
Ref. | Enticott, J. and D. Tipling, 1997 |
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Associations | colony-forming/symphorism; |
Associated with | |
Association remarks | Dense breeding colonies and feeding flocks. Also in Ref. 91362. |
Parasitism |
Feeding type | mainly animals (troph. 2.8 and up) |
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Feeding type Ref. | Karpouzi, V.S., 2005 |
Feeding habit | hunting macrofauna (predator) |
Feeding habit Ref. | Harrison, P., 1987 |
Estimation method | original sample | unfished population | Remark | ||
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Troph | s.e. | Troph | s.e. | ||
From diet composition | |||||
From individual food items | 4.15 | 0.60 | Trophic level estimated from a number of food items using a randomized resampling routine. | ||
Ref. |